Everyone on staff at Tottney Castle knows that the lovely Lady Alyce Marshmorton (Joan Fontaine) must marry soon, so a wager is proposed as to the identity of the lucky man. With all the likely candidates already claimed, young footman Albert (Harry Watson) places a bet on a "Mr. X," someone totally out of the blue.
Lady Alyce secretly has a romantic interest in an American no one from her family has yet met. She leaves the castle one day to venture into London, where by chance she encounters Jerry Halliday (Fred Astaire). He is an American entertainer, accompanied by press agent George (George Burns) and secretary Gracie (Gracie Allen), but he is not well enough known to be recognized by Lady Alyce.
Jerry is incorrectly led to believe that he is the American that Lady Alyce is in love with. He goes to the castle, encouraged by Albert but discouraged by Keggs (Reginald Gardiner), a scheming butler whose money is on another beau. The closest Jerry can get to Lady Alyce is a castle tour, at least until Albert can sneak him upstairs.
False impressions abound, as Jerry also fails to recognize Lady Alyce's father (Montagu Love), the lord of the manor. He is slapped in the face in a Tunnel of Love, misunderstanding the young lady's intentions entirely. In the end, however, he and Lady Alyce do find romance.
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